Pathogenesis and Control of Chronic and Emerging Infections, Universite de Montpellier, National Institute of Health and Medical Research, Montpellier, France ; Unité des Virus Émergents, Aix-Marseille Univ, Università di Corsica, IRD 190, National Institute of Health and Medical Research 1207, Institut de Recherche Biomédicale des Armées, Marseille